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slow-adapting receptors, while phasic receptors are                         Chapter 15  Sensory Pathways and the Somatic Nervous System   555
      fast-adapting receptors. (Figure 15–3)
                                                                                         sensory information from organs between the diaphragm and
15-3  	 General sensory receptors are classified by the type of                          the pelvic cavity. Dorsal roots of spinal nerves S2–S4 carry
                                                                                         sensory information from more inferior structures.
          stimulus that excites them p. 539
	7.	Three types of nociceptors in the body provide information                    15-5  	 The somatic nervous system is an efferent division that

      on pain as related to extremes of temperature, mechanical                              controls skeletal muscles p. 549
      damage, and dissolved chemicals. Myelinated Type A fibers                   	15.	 Somatic motor (descending) pathways always involve an upper
      carry fast pain. Slower Type C fibers carry slow pain.
	8.	Thermoreceptors are found in the dermis. Mechanorecep-                               motor neuron (whose cell body lies in a CNS processing cen-
      tors are sensitive to distortion of their membranes, and in-                       ter) and a lower motor neuron (whose cell body is located in
      clude tactile receptors, baroreceptors, and proprioceptors.                        a nucleus of the brain stem or spinal cord). (Figure 15–8)
      There are six types of tactile receptors in the skin, and three             	16.	 The neurons of the primary motor cortex are pyramidal cells.
      kinds of proprioceptors. Chemoreceptors include carotid                            The corticospinal pathway provides voluntary skeletal muscle
      bodies and aortic bodies. (Figure 15–4)                                            control. The corticobulbar tracts end at the cranial nerve nu-
                                                                                         clei. The corticospinal tracts synapse on lower motor neurons
15-4  	 Separate pathways carry somatic sensory and visceral                             in the anterior gray horns of the spinal cord. The corticospinal
                                                                                         tracts are visible along the medulla oblongata as a pair of thick
          sensory information p. 544                                                     bands, the pyramids, where most of the axons decussate to
	9.	Sensory neurons that deliver sensation to the CNS are referred                       enter the descending lateral corticospinal tracts. Those that
                                                                                         do not cross over enter the anterior corticospinal tracts. The
      to as first-order neurons. These synapse on second-order                           corticospinal pathway provides a rapid, direct mechanism for
      neurons in the brain stem or spinal cord. The next neuron in                       controlling skeletal muscles. (Figure 15–9; Table 15–2)
      this chain is a third-order neuron, found in the thalamus.                  	17.	 The medial and lateral pathways include several other cen-
	10.	 Three major somatic sensory pathways carry sensory informa-                        ters that issue motor commands as a result of processing per-
      tion from the skin and musculature of the body wall, head,                         formed at a subconscious level. (Table 15–2)
      neck, and limbs: the spinothalamic pathway, the posterior column
      pathway, and the spinocerebellar pathway. (Figure 15–5)                     	15	18.	 The medial pathway primarily controls gross movements of
	11.	 The spinothalamic pathway carries poorly localized                                 the neck, trunk, and proximal limbs. It includes the vestibulo-
      (“crude”) sensations of touch, pressure, pain, and temperature.                    spinal, tectospinal, and reticulospinal tracts. The vestibulospi-
      The axons involved decussate in the spinal cord and ascend                         nal tracts carry information related to maintaining balance
      within the anterior and lateral spinothalamic tracts to                            and posture. Commands carried by the tectospinal tracts
      the ventral nuclei of the thalamus. Abnormalities along the                        change the position of the head, neck, and upper limbs in
      spinothalamic pathway can lead to phantom limb pain,                               response to bright lights, sudden movements, or loud noises.
      painful sensations that are perceived as real, and referred                        Motor commands carried by the reticulospinal tracts vary
      pain, inaccurate localizations of the source of pain.                              according to the region stimulated. (Table 15–2)
      (Spotlight Figure 15–6, Figure 15–7; Table 15–1)                            	19.	 The lateral pathway consists of the rubrospinal tracts, which
	12.	 The posterior column pathway carries fine touch, pressure,
      and proprioceptive sensations. The axons ascend within the                         primarily control muscle tone and movements of the distal
      fasciculus gracilis and fasciculus cuneatus and relay infor-                       muscles of the upper limbs. (Table 15–2)
      mation to the thalamus by the medial lemniscus. Before the                  	20.	 The basal nuclei adjust the motor commands issued in other
      axons enter the medial lemniscus, they cross over to the                           processing centers and provide background patterns of move-
      opposite side of the brain stem. This crossing over is called                      ment involved in voluntary motor activities.
      decussation. (Spotlight Figure 15–6; Table 15–1)                            	21.	 The cerebellum monitors proprioceptive sensations, visual in-
	13.	 The spinocerebellar pathway, including the posterior and                           formation, and vestibular (balance) sensations. The integrative
      anterior spinocerebellar tracts, carries sensations to the                         activities performed by neurons in the cortex and nuclei of the
      cerebellum concerning the position of muscles, tendons, and                        cerebellum are essential for the precise control of movements.
      joints. (Spotlight Figure 15–6; Table 15–1)                                 	22.	 Spinal and cranial reflexes provide rapid, involuntary, pre-
	14.	 Visceral sensory pathways carry information collected by in-                       programmed responses that preserve homeostasis. Voluntary
      teroceptors. Sensory information from cranial nerves V, VII,                       responses are more complex and require more time to prepare
      IX, and X is delivered to the solitary nucleus in the medulla                      and execute.
      oblongata. Dorsal roots of spinal nerves T1–L2 carry visceral               	23.	 During development, the spinal and cranial reflexes are first
                                                                                         to appear. Complex reflexes develop over years, as the CNS
                                                                                         matures and the brain grows in size and complexity.

Review Questions	                                                                                         See the blue Answers tab at the back of the book.

LEVEL 1  Reviewing Facts and Terms                                                	 2.	 ___________ receptors are normally inactive, but become active for a
	 1.	 The larger the receptive field, the (a) larger the stimulus needed to              short time whenever there is a change in the modality that they monitor.

       stimulate a sensory receptor, (b) fewer sensory receptors there are,       	 3.	 In sensory organ function what is adaptation?
       (c) harder it is to locate the exact point of stimulation, (d) larger the  	 4.	 The area of primary sensory cortex devoted to a body region is relative
       area of the somatosensory cortex in the brain that deals with the area,
       (e) closer together the receptor cells.                                           to the (a) size of the body area, (b) distance of the body area from the
                                                                                         brain, (c) number of motor units in the area of the body, (d) number
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