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slow-adapting receptors, while phasic receptors are Chapter 15 Sensory Pathways and the Somatic Nervous System 555
fast-adapting receptors. (Figure 15–3)
sensory information from organs between the diaphragm and
15-3 General sensory receptors are classified by the type of the pelvic cavity. Dorsal roots of spinal nerves S2–S4 carry
sensory information from more inferior structures.
stimulus that excites them p. 539
7. Three types of nociceptors in the body provide information 15-5 The somatic nervous system is an efferent division that
on pain as related to extremes of temperature, mechanical controls skeletal muscles p. 549
damage, and dissolved chemicals. Myelinated Type A fibers 15. Somatic motor (descending) pathways always involve an upper
carry fast pain. Slower Type C fibers carry slow pain.
8. Thermoreceptors are found in the dermis. Mechanorecep- motor neuron (whose cell body lies in a CNS processing cen-
tors are sensitive to distortion of their membranes, and in- ter) and a lower motor neuron (whose cell body is located in
clude tactile receptors, baroreceptors, and proprioceptors. a nucleus of the brain stem or spinal cord). (Figure 15–8)
There are six types of tactile receptors in the skin, and three 16. The neurons of the primary motor cortex are pyramidal cells.
kinds of proprioceptors. Chemoreceptors include carotid The corticospinal pathway provides voluntary skeletal muscle
bodies and aortic bodies. (Figure 15–4) control. The corticobulbar tracts end at the cranial nerve nu-
clei. The corticospinal tracts synapse on lower motor neurons
15-4 Separate pathways carry somatic sensory and visceral in the anterior gray horns of the spinal cord. The corticospinal
tracts are visible along the medulla oblongata as a pair of thick
sensory information p. 544 bands, the pyramids, where most of the axons decussate to
9. Sensory neurons that deliver sensation to the CNS are referred enter the descending lateral corticospinal tracts. Those that
do not cross over enter the anterior corticospinal tracts. The
to as first-order neurons. These synapse on second-order corticospinal pathway provides a rapid, direct mechanism for
neurons in the brain stem or spinal cord. The next neuron in controlling skeletal muscles. (Figure 15–9; Table 15–2)
this chain is a third-order neuron, found in the thalamus. 17. The medial and lateral pathways include several other cen-
10. Three major somatic sensory pathways carry sensory informa- ters that issue motor commands as a result of processing per-
tion from the skin and musculature of the body wall, head, formed at a subconscious level. (Table 15–2)
neck, and limbs: the spinothalamic pathway, the posterior column
pathway, and the spinocerebellar pathway. (Figure 15–5) 15 18. The medial pathway primarily controls gross movements of
11. The spinothalamic pathway carries poorly localized the neck, trunk, and proximal limbs. It includes the vestibulo-
(“crude”) sensations of touch, pressure, pain, and temperature. spinal, tectospinal, and reticulospinal tracts. The vestibulospi-
The axons involved decussate in the spinal cord and ascend nal tracts carry information related to maintaining balance
within the anterior and lateral spinothalamic tracts to and posture. Commands carried by the tectospinal tracts
the ventral nuclei of the thalamus. Abnormalities along the change the position of the head, neck, and upper limbs in
spinothalamic pathway can lead to phantom limb pain, response to bright lights, sudden movements, or loud noises.
painful sensations that are perceived as real, and referred Motor commands carried by the reticulospinal tracts vary
pain, inaccurate localizations of the source of pain. according to the region stimulated. (Table 15–2)
(Spotlight Figure 15–6, Figure 15–7; Table 15–1) 19. The lateral pathway consists of the rubrospinal tracts, which
12. The posterior column pathway carries fine touch, pressure,
and proprioceptive sensations. The axons ascend within the primarily control muscle tone and movements of the distal
fasciculus gracilis and fasciculus cuneatus and relay infor- muscles of the upper limbs. (Table 15–2)
mation to the thalamus by the medial lemniscus. Before the 20. The basal nuclei adjust the motor commands issued in other
axons enter the medial lemniscus, they cross over to the processing centers and provide background patterns of move-
opposite side of the brain stem. This crossing over is called ment involved in voluntary motor activities.
decussation. (Spotlight Figure 15–6; Table 15–1) 21. The cerebellum monitors proprioceptive sensations, visual in-
13. The spinocerebellar pathway, including the posterior and formation, and vestibular (balance) sensations. The integrative
anterior spinocerebellar tracts, carries sensations to the activities performed by neurons in the cortex and nuclei of the
cerebellum concerning the position of muscles, tendons, and cerebellum are essential for the precise control of movements.
joints. (Spotlight Figure 15–6; Table 15–1) 22. Spinal and cranial reflexes provide rapid, involuntary, pre-
14. Visceral sensory pathways carry information collected by in- programmed responses that preserve homeostasis. Voluntary
teroceptors. Sensory information from cranial nerves V, VII, responses are more complex and require more time to prepare
IX, and X is delivered to the solitary nucleus in the medulla and execute.
oblongata. Dorsal roots of spinal nerves T1–L2 carry visceral 23. During development, the spinal and cranial reflexes are first
to appear. Complex reflexes develop over years, as the CNS
matures and the brain grows in size and complexity.
Review Questions See the blue Answers tab at the back of the book.
LEVEL 1 Reviewing Facts and Terms 2. ___________ receptors are normally inactive, but become active for a
1. The larger the receptive field, the (a) larger the stimulus needed to short time whenever there is a change in the modality that they monitor.
stimulate a sensory receptor, (b) fewer sensory receptors there are, 3. In sensory organ function what is adaptation?
(c) harder it is to locate the exact point of stimulation, (d) larger the 4. The area of primary sensory cortex devoted to a body region is relative
area of the somatosensory cortex in the brain that deals with the area,
(e) closer together the receptor cells. to the (a) size of the body area, (b) distance of the body area from the
brain, (c) number of motor units in the area of the body, (d) number

