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S p ot l i g h t592  Unit 3  Control and Regulation     Figure 17–2

                                                        Olfaction and Gustation

                                                        Olfaction and gustation are special senses                          Stimulus
                                                        that give us vital information about our                                           Dendrites
                                                        environment. Although the sensory
                                                        information is diverse and complex, each
                                                        special sense originates at receptor cells
                                                        that may be neurons or specialized
                                                        receptor cells that communicate with
                                                        sensory neurons.

      OLFACTION                                                                                Stimulus                     Specialized
                                                                                 Action removed                             olfactory
Olfactory receptors are the dendrites                                          potentials                                   neuron
of specialized neurons. When                            +30
dissolved chemicals contact the                                                                                     –70 mV
dendritic processes, there is a
depolarization, called a generator               mV mV     0                                             Threshold
potential. This graph shows the
action potentials produced by a                                Stimulus
generator potential.
                                                        –60
                                                        –70 Generator potential

                                                        0                                                                   to CNS
                                                                        Time (msec)

      GUSTATION                                                                     RECEPTOR CELL                           Stimulus
                                                        +30
The receptors for the senses of taste, vision,
equilibrium, and hearing are specialized                0                        Stimulus
cells that have unexcitable membranes and                    Stimulus            removed
form synapses with the processes of sensory
neurons. When stimulated, the membrane                  –60 Threshold                                               –70 mV            Receptor
of the receptor cell undergoes a graded                 –70 Depolarization                                          –70 mV            cell
depolarization that triggers the release of                                                                                           Synapse at
chemical transmitters at the synapse. These             0                                                                             dendrite
transmitters then depolarize the sensory                                Time (msec)                                                   Sensory
neuron, creating a generator potential and                                                                                            neuron
action potentials that are propagated to the
CNS. Because a synapse is involved, there is                    Stimulus         AXON
a slight synaptic delay. However, this                  +30                               Action
arrangement permits modification of the
sensitivity of the receptor cell by presynaptic                        Synaptic        potentials
facilitation or inhibition.                                0 delay
                                                 mV
   592
                                                        –60                          Generator potential                    to CNS
                                                        –70                      Time (msec)

                                                                     0
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